Ectillænus giganteus

Here is a trilobite collected in France, famous in Brittany and Spain. This specimen has an unusual transversal curve, moreover when one know that this trilobite is frequently found strongly distorted, when not totally flat. (cf. infra). This specimen does indeed himself suffer a transversal distortion that let it symetrical, bending him transversaly.

We remind that the appearance of trilobites may strongly differ from one specimen to another simply because of tectonic distortions, but this is related to sediments too. The specimes herein reported collected in the ardoisières de La Pouëze are strinking examples of very different conservation modes.

 

Description du spécimen

  • Partially enrolled trilobite..
  • on a piece of matrix.
  • Small right eye.
  • Thoracic rings perfectly preserved, transversally bent (tectonic distortion)
  • Total size : 105 mm (75 mm)
  • Middle Ordovician (Llandeilo).
  • Normandy.
  • France.
  • NB : The anterior part of the cephalon presents cuticular ridges hardly visible on pictures.
Right lateral view.
Dorsal view
Posterior view.

 

DIAGNOSE

 

Order

Corynexochida

KOBAYASHI 1935 Occurrence: Lower Cambrian – Middle Devonian
  • Cephalon: Opisthoparian sutures.
  • Glabella elongate, sides often concave (pestle-shaped).
  • Furrows (when not effaced) typically with splayed arrangement, the hind pair pointing sharply backwards. Anterior pairs tending more and more forward directed. Sometimes furrows pit-like.
  • Cranidial borders often ledgelike.
  • Hypostome conterminant or (in derived forms) impendent.
  • Eyes typically large, in some gently arcuate
  • Thorax: typically with 7-8 segments (but range for order is 2-12, rarely more).
  • Pleural tips often spinose.
  • Pygidium : typically large (isopygous or subisopygous).
  • Variable form.
  • Some spinose.

Suborder

Illænina

JAANUSSON 1959

Occurence : Upper Cambrian Devonian.

  • Cephalon : Typically effaced.
  • Doublure broad.
  • Opisthoparian facial sutures.
  • Sutures distinctly diverging anteriorly.
  • Glabella expanding forwards.
  • Lateral furrows often faint or absent.
  • 4 pairs of glabellar furrows, commonly faint or absent.
  • Extra-axial cephalic muscle impressions (lunette) present.
  • Eyes frequently on the posterior part of the genæ, close to the axial furrows.
  • Terrace lines frequent, particularly at the distal ends of the thoracic exoskeleton and the ventral doublure.
  • Hypostome impendent (conterminant, but hypostome no longer matching anterior glabellar border)
  • Rostral plate limited by sutures, fused in the late species of Panderia.
  • Trilobitation frequently fainting .
  • Thorax : 8-10 segments.
  • Pygidium : Isopygous or subisopygous.
  • Rounded posteriorly.
  • Usually with short axis.

Superfamily

Illænoidea HAWLE & CORDA 1847  
Family Illænidæ HAWLE & CORDA 1847
  • Cephalon : Axial region of cephalon smooth, merging forward into frontal area without boundary.
  • 4 pairs of muscle scars can be observed instead of glabellar and occipital furrows.
  • Glabella strongly convex.
  • Opisthoparian facial sutures.
  • Rostral shield broad (tr.).
  • Hypostoma oval, with elongated body (sag.)
  • Anterior wings of hypostoma broad, quadrangular.
  • Pygidium : Axis and pleural field smooth or with very faint traces of unfurrowed ribs.
  • Surface ornemented with terrace lines or small pits, or both.
  • No tuberculate or granulose ornementation.
Genus Ectillænus (= Wossekia) SALTER 1867
  • Cephalon : Dorsal furrows almost rectilinear.
  • Eyes very small or absent.
  • Post-ocular branches of the facial sutures curved at their posterior end.
  • Subtriangular hypostoma.
  • Anterior border of the hypostoma almost transverse or slightly bent forward in its central part.
  • Anterior wings broad and short.
  • Thorax : 10 segments.
  • Axis well defined.
  • Pygidium : About half of the cephalon's length (sag.)
  • Doublure as long as 30 to 50 % of the pygidium total length (sag.)
Species giganteus BURMEISTER 1843

Large sized isopygous form (up to 30 cm).

  • Cephalon : parabolic.
  • Almost smooth.
  • 5 faint pairs of muscle scars (see below)
  • Dorsal furrows faint, completely fainting exsagittally at the anterior half.
  • Very small eyes, located at the posterior part of the cranidium.
  • Librigenæ narrow and subtriangular.
  • No genal spines.
  • Cephalic doublure broad with concentric terrace lines.
  • Rostral plate of large size, semilunar, with transverse sutures.
  • Hypostoma quadratic with subtriangular body, sharp and covered with terrace lines.
  • Anterior wings broad (tr.), quadrangular.
  • Cephalic angle : 80 to 85°.
  • Thorax : 10 segments
  • Quadratic rings, as wide as the third of the total width.
  • Smooth pleuræ, bearing a geniculation at mid-length.
  • Pygidium : slightly ogival, almost circular.
  • Axis hardly distinct.
  • Pygidial angle : 110 to 120°

 

Complementary pictures :

 

Cephalon, dorsal view
Géniculations à mi-longueur des plèvres.
Terrasses cuticulaires céphaliques.

Scroll over the central picture to see the pleural geniculation

 

- Note the characteristic dorsal furrows on the cephalon and the terrace lines on the broad cephalic doublure.

 

Partial enrollment. Cephalon is on the right of the picture.

 

Details of the thoracic axis. Note the deep perforations.

 

 

Synthethogram of Ectillænus giganteus. (After P. Lebrun, 2001)

 

Discussion and remarks :

Following RÁBANO & GUTIERREZ (1983), one will note that E. bituberosus KNUPFER 1967 and E. convergatus KNUPFER 1967 of Thuringia (Germany) are no more assigned to the genus Ectillænus.

RÁBANO & GUTIERREZ (1983) precised the diagnose of Ectillænus giganteus with measurements that are sadly frequently useless due to tectonic distortions. They consist in the angle measured between 2 sagital tangents (the tangents to the anterior border and to the occipital ring for the cephalic angle, and the tangents to the proximal part of the axis and to the border for the pygidium). They included in the diagnose the description of the muscular impressions that I write down only here (they are almost never seen, being situated in the inner face of the cranidium) :

Ectillænus has been found in Iberic Peninsula, in France, in Great-Britain, in Bugaria, in Morocco, in Central Asia (?) and in Sweden (?).

 

A few other specimens (La Pouëze, Maine-et-Loire, France).

Pieces of cephalothorax. Specimens presenting a strong lateral distortion, flattened.

 

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