Leviceraurus mammilloides HESSIN 1988

 

Uncommon Trilobite from Canada, this trilobite has been described by Willian Hessin in 1988. It is a middle-upper Ordovician Trilobite from the Cobourg Formation, in the Ontario province. He named his finding from the latin levis (bald) and Ceraurus, in reference to reduced prosopon as compared to that of the related genus, Ceraurus.

The similarities with Ceraurus have been underlined by Hessin, as he precises that isolated pygidia and hypostomes of Leviceraurus can easily be confused with those belonging to some species of Ceraurus.

1936 ?Ceraurus sp. 2 COOPER & KINDLE 1936, p. 367, Pl. 53, fig 28 (only).
1988 Leviceraurus mammilloides HESSIN W. A. 1988. p. 89, Pl. 2, fig. 1 - 6, p. 91, Pl. 3, fig. 1 - 4.

 

Specimen description :

  • Almost complete trilobite.
  • Enrolled on matrix
  • Small granulations on the exosqueletton.
  • Total size : 26 x 12 mm
  • middle-upper Ordovician
  • Cobourg Formation.
  • Ontario, Canada

 

Dorsal view of the specimen.
Antero-lateral view.

 

Diagnosis :

 

Order

 

 

 

 

 

 

 

 

 

Phacopida

 

 

 

 

 

 

 

 

 

SALTER 1864

 

 

 

 

 

 

 

 

 

Occurrence: Lower Ordovician (Tremadoc) to Upper Devonian (Famennian).

  • Cephalon: Proparian (Phacopina and Cheirurina), gonatoparian (Calymenina) or opisthoparian (Calymenina),
  • Preglabellar field often very short or absent.
  • 4 or fewer pairs of glabellar furrows.
  • Eyes : when present, schizochroal (Phacopina) or holochroal (Cheirurina and Calymenina).
  • With rostral plates (Calymenina and Cheirurina) or without (some Phacopina).
  • Hypostome conterminant (all suborders) to impendent (some devonian Phacopina).
  • Exosqueletton generally granulous.
  • Thorax: 8 – 19 segments, sometimes distinctly furrowed.
  • Axis sometimes broad (e.g., Homalonotidae).
  • Pygidium : Typically micropygous (most Calymenina and Phacopina), but variable (e.g., subisopygous in Dalmanitoidea and Acastoidea).
  • May be lobed or spiny (e.g., Cheirurina, some Dalmanitoidea, Acastoidea), or smooth-margined, with round or subtriangular outline (e.g., Calymenina, Phacopoidea).

Sub-order

 

 

Cherirurina

 

 

HARRINGTON & LEANZA 1957

 

  • Cephalon: proparian sutures (a few opisthoparian).
  • Glabella usually expanding forwards or barrel-shaped, up to 4 pairs of furrows;.
  • Holochroal eyes, often small.
  • Gena often with fine pitting.
  • With rostral plate.
  • Hypostome conterminant, with anterior wing bearing prominent process.
    Thorax: variable numbers of distinctive segments within Phacopid range of 8-19.
  • Often spine-tipped.
    Pygidium: often highly modified with 2-16 distinct pairs of ribs.
  • Lobed or spinose margin.
    Other: often with tuberculate exoskeleton.

Family

 

Cheiruridae

 

HAWLE & CORDA 1847

 

  • Thorax : Pleurae with pointed or rounded spines.
  • Oblique or transverse pleural furrows (in some genera represented by pits, or effaced).
  • Pygidium : 2 to 4 pairs of pleural ribs and spines.
  • Axial rings few.
  • Other : surface commonly tuberculate, pitted or both.

Sub-family

 

 

 

Cheirurinae

 

HAWLE & CORDA 1847

 

 

  • Cephalon : Glabella expanding forward or with subparallel sides.
  • Preoccipital lateral glabelar furrows generally curved backward, commonly joining occipital furrow.
  • Hypostoma with elongate middle body.
  • Thorax : generally of 11 segments.
  • Proximal part of pleurae with distinct oblique pleural furrows.
  • Pygidium : with 2 or 3 pairs of pleural spines.
  • 2 posterior pairs more or less reduced in some.

Genus

 

Leviceraurus

HESSIN 1988

  • Cephalon : moderatly inflated and nearly parallel-sided glabella.
  • 3 pairs of short glabellar furrows.
  • Width across occipital ring slightly less than one-fourth width of cephalon.
  • Occipital ring broad, about one-fourth length of glabella.
  • Glabella smooth to unaided eyes, but has prosopon of very fine granules and typically four pairs of low, rounded tubercles on medial portion of glabella (often fourth pair and one third pair absent).
  • Eye small, elevated and positionned on mid-point of cheek opposite 2S furrow or 2p lobe.
  • Rostral plate possessing a pair of faint tubercles.
  • Anterior border very narrow, almost indistinguishable from anterior lobe of glabella.
  • Genal spines moderately long, extanding back to opposite 4th or 5th thoracic segment.
  • Pygidium : Triangular in outline.
  • Excluding slender anterior spines which are almost as long as length of thorax.
  • 2 pairs of posterior spines present but very short.

Species

mammilloides

HESSIN 1988

  • As for genus.

Remarks :

The etymology of mammilloides comes from mammillae (nipples, latin) and oides (like, latin), in reference to tubercles on rostral plate.

Discussion :

To help in differential diagnoses, here a few diverging points :

- Ceraurus pleurexanthemus GREEN (type species for Ceraurus), which occurs with Leviceraurus, can generally be recognized by its coarser prosopon.

- Ceraurus possess a glabella which expands forward to the anterior lobe and an inflated anterior border which is clearly separated from an inflated anterior lobe by a relatively deep preglabellar furrow.

- Ceraurus mackenziensis LUDVIGSEN 1979 resemble Leviceraurus in prosopon and some cephalic details. However, these species have slightly coarser granules, the latter species possesses a nearly parallel-sided glabella like Leviceraurus, but differs in the lack of tubercles on glabella, a glabella that is clearly separated from the anterior border, eyes located very close to the posterior border, a cephalon broadly triangular in outline and a pygidium subtriangular in outline (excluding spines) with indistinct axial rings.

 

Complementary pictures :

Leviceraurus mammilloides
Prosopon with very fine granules.
Prosopon with very fine granules.
Faint tubercles on rostral plate.
Leviceraurus mammilloides
 
Prosopon with very fine granules.

 

Synthetogram of Leviceraurus mammilloides after HESSIN 1988

 

References :

COOPER & KINDLE. New brachiopods and trilobites from Upper Ordovician of Percé, Québec. Journal of palaeontology 10: pp. 348 - 372.

HESSIN W. A. 1988. Leviceraurus. A new cheirurine Trilobite from the Cobourg Formation (Middle-Upper Ordovican) Southern Ontario, Canada. Journal of palaeontology 62 (1) : pp. 87 - 93.

 

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